1881–1883 Natural Sciences Tripos in Cambridge; 1884 Medizinstudium in London; 1885–1887 Forschungsaufenthalte in Frankreich, Spanien, Italien und Deutschland; 1887 Fellow of Gonville and Caius College, Cambridge; 1889–1905 Secretary of the Physiological Society; 1893 Fellow of the Royal Society (London); 1895 Holt Professor of Physiology, Liverpool; 1904 Siliman Lectures (The Integrative Action of the Nervous System); 1913 Waynfleete Professor of Physiology, Oxford; 1920–1925 President of the Royal Society (London); 1932 Nobelpreis für Medizin (gemeinsam mit Edgar Douglas Adrian); 1937/38 ›Gifford Lectures‹ (Man on his Nature).